Abies firma in habitat on northern Kyushu, near the city of Fukuoka [Tom Velardi, 2004].
Crown of an old tree in the forest shown above [Tom Velardi, 2004].
Abies firma in habitat on northern Kyushu, near the city of Fukuoka [Tom Velardi, 2004].
Crown of an old tree in the forest shown above [Tom Velardi, 2004].
Abies firma
モミ [Japanese], Momi fir.
Synonymy (Farjon 1998):
Abies × umbellata Liu 1971 (syn: A. umbellata Mayr 1890, A. homolepis var. umbellata Wilson 1916) is the natural hybrid of A. homolepis and A. firma (Aizawa and Iwaizumi 2020). The parent species represent different subsections of Abies section Momi.
Tree to 50 m. tall and 200 cm dbh. Trunk massive, straight, with long branches horizontal or ascending. Crown broadly pyramidal or domed, becoming flat-topped with age. Bark gray, smooth with resin blisters when young, then fissured, corky and darker with age. Branchlets grooved, yellowish-green to gray-brown, glabrous or with fine blackish pubescence. Buds ovoid to conical, to 10 × 5 mm, slightly resinous, with broadly conical scales, grayish-brown, persisting several years. Juvenile leaves bifid, extremely sharp pointed, present at the base of old trees. Leaves pectinate, nearly 2-3 ranked, spreading outwards and upwards, with a distinct V-like parting, grooved above, keeled below, bright dark green above, glaucous-yellow below; stomata usually absent above, in 12-13 lines below; 1.5-3.5 cm long by 2-4 mm. wide, base twisted, apex obtuse, emarginate or notched. Pollen cones cylindrical, yellowish, solitary and pendant in leaf axils, 2.5-3 cm long. Seed cone ovoid-oblong to conical, apex rounded, green with yellow bracts, maturing yellow-brown, 8-15 cm long by 3-5 cm. wide, resinous; bracts erect and slightly exserted, narrow, diamond-shaped. Seed scales 2-2.5 cm long by 2.8-3.2 cm wide, smooth or longitudinally striated. Seed 6-8 mm long, light brown with a 10-15 mm long brownish wing (Silba 1986, Farjon 1990). See García Esteban et al. (2004) for a detailed characterization of the wood anatomy.
The hybrid, Abies × umbellata, strongly resembles A. homolepis but has greenish-yellow seed cones as in A. firma; it mainly occurs at typically at 1000–1100 m elevation in central Honshu (Christian 2021).
Japan: Honshu, Shikoku, Kyushu, and Yakushima at (50-)300-1000(-1900) m, rarely in pure stands on dry sites but usually in mixed forest with a variety of other conifers (Abies homolepis, Chamaecyparis obtusa, Cryptomeria japonica, Pinus parviflora, Pinus densiflora, Pseudotsuga japonica, Sciadopitys verticillata, Torreya nucifera Tsuga sieboldii) and hardwoods (Carpinus laxifolia, Castanea crenata, Fagus crenata, Fagus japonica, Quercus spp.). Climate warm-temperate to cool, with precipitation >1000 mm (Silba 1986, Farjon 1990, Vidakovic 1991). Hardy to Zone 6 (cold hardiness limit between -23.2°C and -17.8°C) (Bannister and Neuner 2001).
Suzuki (1997) studied this species on Yakushima Island. He cites his own previous work (Suzuki and Susukida 1989) in having found a tree stump with 624 rings on Yakushima. He also cites Nakao (1985) as having derived a ring-counted age of 410 years on Mt. Gozen, Yame, Hukuoka, Japan (33.167°N, 130.833°E).
In Japan, the wood is used for construction and coffins. As with most species having wood of a uniform light color, the Japanese find it exceptionally pleasing. In the U.S. and Europe it is a popular ornamental species.
Tom Velardi (e-mail 2004.04.25) recommends the forest pictured at right, near Fukuoka on Kyushu, on a holy mountain named Homanzan, very near the famous Tenmangu Shrine in the town of Dazaifu. The forest here is old and complex, with a multi-storied composition of mixed deciduous/evergreen broadleaf trees and conifers. The broadleaf trees represent many genera, such asQuercus, Myrica, Camellia, Castenea, Acer, Carpinus, Magnolia, Betula, Tillia, Rhus, and Diospyros).
Aizawa, M. and M. G. Iwaizumi. 2020. Natural hybridization and introgression of Abies firma and Abies homolepis along the altitudinal gradient and genetic insights into the origin of Abies umbellata. Plant Species Biology 35(2):147-157.
Christian, T. 2021. Abies × umbellata from Trees and Shrubs Online, accessed 2022.12.07.
Farjon, Aljos. 1990. Pinaceae: drawings and descriptions of the genera Abies, Cedrus, Pseudolarix, Keteleeria, Nothotsuga, Tsuga, Cathaya, Pseudotsuga, Larix and Picea. Königstein: Koeltz Scientific Books.
Liu Tang-Shui. 1971. A Monograph of the Genus Abies. Taipei: National Taiwan University.
Suzuki E. and Susukida J. 1989. Age structure and regeneration process of temperate coniferous stands in the Segire River basin, Yakushima Island. Japanese Journal of Ecology 39: 45-51 [in Japanese with English summary].
Ando T., Chiba K., Nishimura T., and Tanimoto T. 1977. Temperate fir and hemlock forests in Shikoku. In: Shidei T. and Kira T. (eds.). Primary productivity of Japanese forests. JIB synthesis. Tokyo, Japan: University of Tokyo Press, 16:213-224.
Elwes and Henry 1906-1913 at the Biodiversity Heritage Library (Photos). Elwes and Henry correctly inferred the existence of the hybrid A. × umbellata. This series of volumes, privately printed, provides some of the most engaging descriptions of conifers ever published. Although they only treat species cultivated in the U.K. and Ireland, and the taxonomy is a bit dated, still these accounts are thorough, treating such topics as species description, range, varieties, exceptionally old or tall specimens, remarkable trees, and cultivation. Despite being over a century old, they are generally accurate, and are illustrated with some remarkable photographs and lithographs.
Kurata S. 1971. Illustrated important forest trees of Japan. Vols. 1 and 2. Tokyo, Japan: Chikyu Shuppan Co. Ltd.
Suzuki E. and Tsukahara J. 1987. Age structure and regeneration of old growth Cryptomeria japonica forests on Yakushima Island. Bot. Mag. Tokyo 100:223-241.
Last Modified 2024-11-27
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